During the breeding season, wolves use urine marking as a form of communication to convey several important pieces of information:

1. Territorial Boundaries: Urine marking helps to establish and reinforce the boundaries of a pack's territory. This is crucial during the breeding season when competition for resources and mates intensifies. By marking their territory, wolves signal to other packs to stay away.

2. Reproductive Status: Urine can convey information about the reproductive status of the wolves, particularly the females. Female wolves in estrus may have specific pheromones in their urine that signal their readiness to mate. This information is crucial for male wolves in determining when to approach a female for mating.

3. Identity and Social Status: Urine markings can also convey information about the individual wolf's identity and social status within the pack. This helps maintain the social structure and hierarchy, which is particularly important during the breeding season when tensions can be high.

4. Health and Condition: The scent of urine can provide information about the health and physical condition of a wolf. This can be important for potential mates assessing the fitness of a partner.

Overall, urine marking is a complex form of communication that plays a vital role in the social and reproductive dynamics of wolf packs during the breeding season.